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Beschreibung

Specialist reference and practical guidebook on goat reproductive health, emphasizing reproductive diseases, their clinical management, and production management

Elements of Reproduction and Reproductive Diseases of Goats discusses the reproductive system and various reproductive diseases of goats, with coverage of pathogenesis of diseases, disease prevention, diagnosis, and treatment via drugs and other methods, along with general best management of goats. To aid in reader comprehension and practical application, the book includes colored figures, lined figures, and tables visualizing key concepts.

The book explains reproductive anatomy of both males and females in a systematic way. The main topics in this book include breeding characteristics, pregnancy detection, diagnosis, and treatment, management of infertility, obstetrics, abortion, surgery of the reproductive tract, and care and management of kids. The book emphasizes state-of-the-art research on the physiological and biochemical mechanisms in regulation of reproduction.

Edited by a highly qualified practitioner and contributed to by a wide variety of contributors, each with specialized knowledge in their respective area of knowledge, Elements of Reproduction and Reproductive Diseases of Goats covers sample topics such as:

  • Puberty and sexual maturity, selection criteria, nutrition, parasitic infection, seasonal perspectives, and estrus synchronization
  • Preparation for breeding season, gestation and parturition, lactation, reproductive failures, and postpartum care
  • Diseases of the glans penis and prepuce, urethra, scrotum and inguinal lymph nodes, prostate, and testis
  • Infectious abortive diseases, including chlamydiosis, toxoplasmosis, q fever, brucellosis, and campylobacteriosis, along with non-infectious abortive diseases
  • Ketosis and pregnancy toxemia, hypocalcemia, uterine and rectal prolapse, retained placenta and fetal membranes, metritis and endometritis, pyometra, and mastitis

Elements of Reproduction and Reproductive Diseases of Goats is a highly comprehensive resource on the subject ideal for veterinary practitioners, small ruminant researchers, veterinary students, farm managers, industrialists, and all professionals involved in the raising, care, and breeding of goats, along with students and instructors in related programs of study.

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Veröffentlichungsjahr: 2024

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Table of Contents

Cover

Table of Contents

Title Page

Copyright Page

List of Contributors

Preface

Acknowledgment

Section 1: Goat Reproductive Health

1 Anatomy of the Reproductive System in the Buck and Doe

1.1 Introduction

1.2 Male Reproductive System

1.3 Female Reproductive System

Acknowledgment

References

2 Puberty and Sexual Maturity in the Buck and Doe

2.1 Puberty

2.2 Seasonal Breeding

2.3 Puberty in Bucks

2.4 Puberty in Does

2.5 Estrous Cycle

2.6 Pregnancy

2.7 Measures to Increase Reproductive Performance in Goats

References

Further Reading

3 Reproductive Biology of the Buck and Doe

3.1 Introduction

3.2 Reproduction in Goats

3.3 Male Reproductive Tract

3.4 Female Reproductive System

3.5 Seasonal Sexual Activity

3.6 Estrous Cycle

3.7 Pregnancy

3.8 Kidding

3.9 Lactation

3.10 Female with Newborn Kids

3.11 Control of Sexual Activity and Optimization of Fertility Results

3.12 Estrus and Male Mating Behavior

References

Further Reading

4 Selection Criteria for the Buck and Doe

4.1 Introduction

4.2 Why Selection is Important

4.3 Setting a Breeding Objective

4.4 Breeding and Selection Tools

Further Reading

5 Impact of Nutrition on Reproduction in Goats

5.1 Introduction

5.2 Goat Feeding

5.3 Nutritional Health Problems

5.4 Conclusion

References

6 Effect of Parasitic Infection on Reproductive Health

6.1 Introduction

6.2 Parasitic Diseases and Reproductive Health

6.3 Conclusion

References

7 Seasonal Perspectives on Reproduction of the Buck and Doe

7.1 Introduction

7.2 Influence of Photoperiod on Reproductive Seasonality

7.3 Circadian Rhythm and Reproduction in Mammals

7.4 Other Factors That Influence Reproductive Seasonality

7.5 Melatonin

7.6 Control of Sexual Activity

References

8 The Buck Effect: A Management Tool for Improving Reproductive Efficiency in Goats

8.1 Introduction

8.2 The Buck Effect

8.3 Synchronization of Estrus by Hormonal Protocol

8.4 Behavior of Goats During Estrus

8.5 Hormones Related to Estrus and Their Changes in Goats

8.6 Conclusion

References

9 Estrus Synchronization and Mating Systems in Goats

9.1 Introduction

9.2 Estrous Cycle

9.3 The Male Effect

9.4 Applications of Synchronization of Estrus

9.5 Female Mating Systems

References

10 Preparation for the Breeding Season

10.1 Purpose of Goat Breeding

10.2 Establishing a Reproducing/Breeding Flock

10.3 Examination for Breeding Soundness in Goats

10.4 Onset of Puberty

10.5 Breeding During Puberty

10.6 Preparation of Males for Breeding

10.7 Preparation of Females for Breeding

10.8 Estrous Cycle

10.9 Goat Nutrition During Breeding

10.10 Breeding Systems

10.11 Mating Methods

10.12 Goat Gestation Period

10.13 Pregnancy Diagnosis

10.14 Separating Male and Female Goats

References

11 Gestation and Parturition

11.1 Introduction

11.2 Gestation

11.3 Parturition or Kidding

11.4 Problems of Caprine Gestation

11.5 Pharmacological Induction of Kidding

References

12 Lactation

12.1 Introduction

12.2 Mammary Glands are Sophisticated Sweat Glands

12.3 Colostrogenesis

12.4 Lactogenesis

12.5 Involution of Mammary Gland

12.6 Lactation Curve

12.7 Heat Stress

12.8 Dry Period for Cell Proliferation

12.9 Conclusion

References

13 Reproductive Techniques

13.1 Introduction

13.2 Reproductive System of the Doe

13.3 Reproductive Techniques in Goats

References

14 Reproductive Failure in Does

14.1 Introduction

14.2 Ovarian Dysfunction

14.3 Disorders of Fertilization

14.4 Pregnancy Wastage

14.5 Perinatal and Neonatal Mortality

15 Postpartum Care of the Doe

15.1 Introduction

15.2 The Puerperium

15.3 Factors Influencing the Puerperium

References

Section 2: Disorders of the Male Reproductive System

16 Diseases of the Glans Penis and Prepuce

16.1 Introduction

16.2 Comparative Anatomy of the Glans Penis

16.3 Anatomy of the Glans Penis in Goats

16.4 Physiology of the Glans Penis and Prepuce in Goats

16.5 Diseases of the Glans Penis and Prepuce in Goats

16.6 Trauma and Injuries

16.7 Conclusion

References

17 Diseases of the Urethra

17.1 Introduction

17.2 General Function of the Goat's Urethra

17.3 Anatomy and Function of the Male Urethra

17.4 Anatomy and Function of the Female Urethra

17.5 Diseases of the Urethra

17.6 General Management and Prevention

17.7 Conclusion

References

18 Diseases of the Inguinal Lymph Nodes

18.1 Introduction

18.2 Lymphatic System Overview

18.3 Role of Inguinal Lymph Nodes

18.4 Location and Structure

18.5 Immune Function

18.6 Common Infectious Diseases Affecting Inguinal Lymph Nodes in Goats

18.7 Noninfectious Causes of Inguinal Lymph Node Enlargement

18.8 Diagnosis of Inguinal Lymph Node Diseases

18.9 Treatment Strategies

18.10 Prevention and Control Measures

18.11 Conclusion

References

19 Diseases of the Prostate

19.1 Introduction

19.2 Cell Biology

19.3 Prostatic Secretions

19.4 Diseases of the Prostrate

19.5 Conclusion

References

20 Diseases of the Scrotum

20.1 Introduction

20.2 Structure and Anatomy

20.3 Diseases of the Scrotum of Bucks

20.4 Conclusion

References

21 Diseases of the Testes

21.1 Introduction

21.2 Epididymis

21.3 Ductus Deferens

21.4 Effect of Season on Buck Puberty and Reproduction

21.5 Diseases of the Testes

21.6 Infectious Diseases Causing Orchitis and Epididymitis

References

Section 3: Diseases of Pregnancy

22 Abortion: General Considerations

22.1 Introduction

22.2 Common Causes of Abortion

22.3 Noninfectious Causes of Abortion

22.4 Infectious Causes of Abortion

22.5 Conclusion

References

23 Infectious Abortive Diseases: Chlamydiosis

23.1 Introduction

23.2 Etiology

23.3 Epidemiology

23.4 Developmental Cycle

23.5 Transmission

23.6 Pathogenesis

23.7 Clinical Signs

23.8 Histopathology

23.9 Differential Diagnosis

23.10 Laboratory Diagnosis

23.11 Prevention and Control

23.12 Economic and Zoonotic Importance

References

24 Infectious Abortive Diseases: Toxoplasmosis

24.1 Introduction

24.2 History

24.3 Taxonomy

24.4 Epidemiology

24.5 Toxoplasma gondii: An Apicomplexan Parasite

24.6 Pathogenesis

24.7 Clinical Signs

24.8 Pathological Findings

24.9 Differential Diagnosis

24.10 Diagnosis

24.11 Treatment

24.12 Prevention and Control

24.13 Public Health Importance

References

25 Infective Abortive Diseases: Q Fever

25.1 Introduction

25.2 Transmission and Spread

25.3 Routes of Infection

25.4 Pathogenesis

25.5 Clinical Signs

25.6 Diagnosis

25.7 Control Methods and Vaccination

25.8 Perspectives for the Future

25.9 Conclusion

References

26 Infectious Abortive Diseases: Brucellosis

26.1 Introduction

26.2 Etiology

26.3 Geographical Distribution

26.4 Zoonotic Importance

26.5 Transmission of B. melitensis

26.6 Pathogenesis and Immune Response

26.7 Clinical Signs

26.8 Postmortem Lesions

26.9 Diagnosis

26.10 Biomarkers of Brucella and Brucellosis

26.11 Treatment

26.12 Control

References

27 Infectious Abortive Diseases: Campylobacteriosis

27.1 Etiology

27.2 The Genus Campylobacter

27.3 Detection, Isolation, and Confirmation

27.4 Clinical Signs

27.5 Pathology

27.6 Epidemiology and Transmission

27.7 Occurrence and Distribution

27.8 Diagnostic Tests and Specimens

27.9 Zoonotic Potential

27.10 Campylobacteriosis and Abortion Storms in Goats

27.11 Treatment, Prevention, and Control

References

28 Noninfectious Abortive Diseases: Toxic Plants, Nutritional, and Genetic

28.1 Introduction

28.2 Toxic Plants

28.3 Nutritional Effects on Reproduction

28.4 Genetic Effects

28.5 Conclusion

References

29 Management of Abortive Diseases

29.1 Introduction

29.2 Causes of Abortion in Goats

29.3 General Approach for the Management of Abortion in Goats

29.4 Conclusion

References

Section 4: Metabolic Diseases

30 Ketosis/Pregnancy Toxemia

30.1 Introduction

30.2 Etiology

30.3 Risk Factors for Pregnancy Toxemia in Goats

30.4 Risk Factors for Ketosis in Goats

30.5 Epidemiology

30.6 Pathophysiology

30.7 Clinical Findings

30.8 Diagnosis

30.9 Treatment

30.10 Prevention

30.11 Conclusion

References

31 Hypocalcemia

31.1 Introduction

31.2 Calcium in Goats

31.3 Regulation of Calcium Homeostasis

31.4 Factors Influencing Absorption of Calcium

31.5 Calcium Deficiencies

31.6 Epidemiology of Hypocalcemia in Goats

31.7 Predisposing Factors for Hypocalcemia in Goats

31.8 Etiology for Hypocalcemia in Goats

31.9 Pathogenesis of Hypocalcemia in Goats

31.10 Clinical Signs of Hypocalcemia in Goats

31.11 Diagnosis of Hypocalcemia in Goats

31.12 Differential Diagnosis of Hypocalcemia in Goats

31.13 Treatment of Hypocalcemia in Goats

31.14 Prevention and Control

References

Section 5: Postpartum Disorders

32 Uterine and Rectal Prolapse

32.1 Uterine Prolapse

32.2 Rectal Prolapse

References

33 Retained Placenta and Fetal Membranes

33.1 Introduction

33.2 Structural Components of Placenta

33.3 Basic Features of Placenta

33.4 Physiology of Expulsion of Placenta

33.5 Clinical Signs

33.6 Therapeutic Management

33.7 Discussion

33.8 Preventive Measures

33.9 Conclusion

References

34 Metritis

34.1 Introduction

34.2 Definition

34.3 Diagnosis

34.4 Treatment

34.5 Conclusion

References

35 Pyometra

35.1 Introduction

35.2 Incidence

35.3 Pathogenesis of Postpartum Uterine Infection

35.4 Etiology

35.5 Clinical Signs

35.6 Diagnosis

35.7 Prognosis

35.8 Treatment

35.9 Managing Female Goats to Prevent Uterine Diseases

35.10 Conclusion

References

36 Mastitis

36.1 Introduction

36.2 Etiology

36.3 Udder Immunity

36.4 Risk Factors Associated with Mastitis

36.5 Clinical Signs

36.6 Diagnosis

36.7 Treatment and Control

References

37 General Preparations for the Newborn Kid

37.1 Introduction

37.2 Care of Newborn Kids

37.3 General Preparations to Ensure the Kid's Well‐being

37.4 Conclusion

References

Further Reading

38 Feeding of Kids

38.1 Introduction

38.2 Colostrum – Gold Standard for Goat Kids

38.3 Tube Feeding in Kids

The following steps should be taken to tube feed a kid (Smith 2021)

38.4 General Kid Nutrition

38.5 Conclusion

References

Further Reading

Section 6: Early Diseases of Kids

39 Stillborn, Weak or Dead Kid

39.1 Introduction

39.2 Biological Variables that Affect Lamb and Kid Survival

39.3 Neonatal Survival

39.4 Early Embryonic Death vs Abortion vs Stillbirth

39.5 General Patterns of Kid Survival from Birth and Post Weaning

39.6 Conclusion

References

40 Hypothermia and Hypoglycemia

40.1 Introduction

40.2 Hypothermia

40.3 Hypoglycemia

References

41 Bacteria and Toxins

41.1 Introduction

41.2 Bacillus anthracis

41.3 Clostridium Species

41.4 Corynebacterium pseudotuberculosis

41.5 Escherichia coli

41.6 Fusobacterium necrophorum

41.7 Listeria monocytogenes

41.8 Mannheimia haemolytica

41.9 Staphylococcus aureus

41.10 Trueperella pyogenes

41.11 Conclusion

References

42 Pneumonia

42.1 Introduction

42.2 Classification of Pneumonia

42.3 Bronchopneumonia

42.4 Interstitial Pneumonia

42.5 Embolic Pneumonia

42.6 Granulomatous Pneumonia

42.7 Transmission Routes

42.8 Clinical Signs

42.9 Diagnosis

42.10 Treatment

42.11 Prevention and Control

References

43 Scours (Diarrhea)

43.1 Introduction

43.2 Etiology

43.3 Infectious Agents Causing Scours

43.4 Noninfectious Agents Causing Scours

43.5 Conclusion

References

44 Navel and Joint Ill

44.1 Introduction

44.2 Navel Ill

44.3 Joint Ill

44.4 Conclusion

References

Section 7: Late Diseases of Kids

45 Coccidiosis

45.1 Introduction

45.2 Classification

45.3 Epidemiology and Immunology

45.4 Life Cycle and Mode of Transmission

45.5 Pathogenesis

45.6 Clinical Signs

45.7 Postmortem Lesions

45.8 Diagnosis

45.9 Differential Diagnosis

45.10 Treatment

45.11 Prevention

45.12 Conclusion

References

46 White Muscle Disease

46.1 Introduction

46.2 Pathophysiology

46.3 Additional Factors Contributing White Muscle Disease

46.4 Clinical Signs

46.5 Diagnosis

46.6 Prevention

46.7 Control

46.8 Conclusion

References

47 Enterotoxemia

47.1 Introduction

47.2 Etiology

47.3 Epidemiology

47.4 Economic Impact

47.5 Pathogenesis

47.6 Clinical Signs

47.7 Diagnosis

47.8 Treatment and Control

47.9 Conclusion

References

48 Acidosis

48.1 Introduction

48.2 Subacute Ruminal Acidosis

48.3 Etiology

48.4 Predisposing Factors

48.5 Pathogenesis

48.6 Clinical Signs

48.7 Diagnosis

48.8 Treatment

48.9 Complications

48.10 Preventive Measures

48.11 Conclusion

References

49 Urolithiasis

49.1 Introduction

49.2 Etiology

49.3 Clinical Signs

49.4 Diagnosis

49.5 Pathology

49.6 Treatment

49.7 Prevention

References

50 Polioencephalomalacia

50.1 Introduction

50.2 Etiology

50.3 Clinical Signs

50.4 Pathology

50.5 Diagnosis

50.6 Treatment

50.7 Prevention

References

51 Salmonellosis

51.1 Introduction

51.2 Etiology

51.3 Distribution

51.4 Transmission

51.5 Clinical Signs of Salmonellosis in Goats and Humans

51.6 Diagnosis

51.7 Treatment

51.8 Prevention and Control

References

52 Rectal Prolapse in Kids

52.1 Introduction

52.2 Prevalence and Etiology

52.3 Classification of Rectal Prolapse

52.4 Pathophysiology

52.5 Treatment

52.6 Aftercare and Complications

52.7 Conclusion

References

53 Copper Deficiency

53.1 Introduction

53.2 Effects of Copper Deficiency on Goat Reproductive Performance

53.3 Signs and Diagnosis of Copper Deficiency in Goats

53.4 Copper Supplementation and Management

53.5 Conclusion

References

Index

End User License Agreement

List of Tables

Chapter 3

Table 3.1 Reproductive organs and functions of male goats.

Table 3.2 Reproductive organs and functions of female goats.

Chapter 4

Table 4.1 Commonly used terms for male and female goats.

Table 4.2 Kid preweaning and weaning weight adjustment factors.

Chapter 5

Table 5.1 Small breeds (slaughter weight 20–25 kg).

Table 5.2 Large breeds (slaughter weight 30–35 kg).

Table 5.3 Doe: nonlactating pregnant (first 15 weeks of gestation).

Table 5.4 Doe: nonlactating pregnant (last six weeks of gestation).

Table 5.5 Doe: lactating (first half of lactation period).

Table 5.6 Doe: lactating (latter half of lactation period).

Table 5.7 Bucks (breeding adult and yearling).

Table 5.8 Cheap concentrate mixture (percentage).

Table 5.9 Mineral mixture for goats.

Table 5.10 Daily nutrient requirements of goats (maintenance, growth, and ea...

Table 5.11 Starter/creep rations.

Table 5.12 Concentrate mixtures for lactating goats.

Table 5.13 Nutrient requirements of goats for general feeding practice.

Table 5.14 Acceptable quantities of macro‐ and microminerals in a goat's die...

Chapter 9

Table 9.1 Species variation in estrous cycle characteristics.

Table 9.2 Timeline of different agents used for synchronization of estrus i...

Table 9.3 How AI will be helpful.

Chapter 11

Table 11.1 Time of production and agents of maternal recognition of pregnan...

Table 11.2 Gestation length in different goat breeds.

Table 11.3 Reproductive traits associated with gestation and parturition in...

Chapter 14

Table 14.1 Causes of embryonic mortality.

Table 14.2 Noninfectious causes of pregnancy wastage in farm animals.

Table 14.3 Summary of diseases causing pregnancy wastage in sheep and goats...

Chapter 22

Table 22.1 Common causes of abortion.

Table 22.2 Noninfectious causes of abortion.

Table 22.3 Minerals and their deficiency symptoms.

Table 22.4 Effect of toxic plant ingestion during pregnancy in goats.

Table 22.5 Infectious causes of abortion and their symptoms.

Chapter 24

Table 24.1 Treatment of toxoplasmosis in small ruminants.

Chapter 32

Table 32.1 Clinical features of uterine prolapse in goats.

Table 32.2 Classification of rectal prolapse by tissue layer.

Table 32.3 Classification of rectal prolapse by severity.

Chapter 36

Table 36.1 Different groups of pathogens as causative agents of caprine mas...

Table 36.2 Scores from the California Mastitis Test on goat milk.

Table 36.3 Interpretation of somatic cell count (SCC) from individual goat ...

Chapter 37

Table 37.1 The process of kidding in goats.

Table 37.2 Feeding of goat kids (0–90 days).

Table 37.3 Vaccination schedule for goats.

Table 37.4 Deworming in goat kids.

Chapter 38

Table 38.1 Method of feeding milk replacer diet.

Table 38.2 Feeding of goat kids (0–90 days).

Table 38.3 Compositions of various creep mixtures for goat kids.

Table 38.4 Composition of concentrate mixture for growing kids (3–12 months...

Table 38.5 Feeding schedule of growing kids 3–12 months.

Chapter 41

Table 41.1 Details of toxins produced by bacterial organisms affecting goat...

Chapter 43

Table 43.1 List of infectious pathogens of scours with zoonotic potential....

Chapter 45

Table 45.1 Valid Eimeria species in goats.

Table 45.2 Merogony, gametogony, prepatent and patent period of different s...

Chapter 47

Table 47.1 Clinical signs in goats affected by enterotoxemia.

Chapter 50

Table 50.1 Main causes of polioencephalomalacia in ruminants.

Table 50.2 Some thiaminase‐containing plant species (Cebra and Cebra 2004; ...

Table 50.3 Some thiamine antagonists (Tylicki et al. 2018; Hrubša et al. 20...

Table 50.4 Some sulfur‐rich dietary ingredients (Cebra and Cebra 2004; Cebr...

Chapter 52

Table 52.1 Classification of rectal prolapse based on rectal tissue involve...

Table 52.2 Classification of rectal prolapse based on rectal layer involved...

Chapter 53

Table 53.1 Effects of copper deficiency on goat reproductive performance.

Table 53.2 Mechanisms and effects of copper deficiency on goat reproductive...

Table 53.3 Laboratory tests for copper deficiency in goats.

Table 53.4 Recommended copper supplementation for goats.

List of Illustrations

Chapter 1

Figure 1.1 Diagrammatic presentation of the male reproductive system.

Figure 1.2 Photograph showing the organs of the male reproductive system.

Figure 1.3 Photograph showing the testis and tunica vaginalis.

Figure 1.4 Photograph (sagittal section) showing the septula testis.

Figure 1.5 Diagrammatic presentation of the female reproductive system.

Figure 1.6 Photograph showing the organs of the female reproductive system....

Chapter 2

Figure 2.1 Schematic representation of the different physiological events oc...

Figure 2.2 Schematic representation of the estrous cycle of the goat.

Figure 2.3 Evolution of sexual behavior during the estrous cycle.

Chapter 3

Figure 3.1 Anatomy of the buck's reproductive tract.

Figure 3.2 Anatomy of the doe's reproductive tract.

Figure 3.3 Buck exhibiting the curled lip and wrinkled nose typical of the F...

Chapter 4

Figure 4.1 Body parts of the goat. 1 Tail. 2 Hip. 3 Rump. 4 Loin. 5 Rib. 6 B...

Figure 4.2 The importance of selection.

Figure 4.3 A healthy buck.

Figure 4.4 Dentition of the goat.

Figure 4.5 The scrotum and its measurement.

Figure 4.6 A goat flock.

Figure 4.7 Udder of a doe.

Chapter 5

Figure 5.1 Feeding of poor‐quality feed.

Figure 5.2 Feeding of growing goats.

Figure 5.3 Feeding of lactating goats.

Figure 5.4 Grazing of bucks.

Figure 5.5 Pasture for feeding of goats.

Figure 5.6 Feeding of concentrates.

Chapter 6

Figure 6.1 Effect of parasitic disease on reproduction in goats.

Figure 6.2 Parasitic causes of reproductive failure in goats.

Figure 6.3 Pale mucous membrane in a hemonchosis affected goat.

Figure 6.4 Weak and emaciated goat due to diarrhea caused by parasitic nemat...

Figure 6.5 Diarrhea due to gastrointestinal helminths in a goat.

Figure 6.6 Abortion in a goat.

Figure 6.7 Strategies for control of parasitic diseases in goats.

Chapter 7

Figure 7.1 Photoperiod and reproduction.

Figure 7.2 Synthesis of melatonin.

Chapter 8

Figure 8.1 Estrus‐related behaviors in goats.

Chapter 9

Figure 9.1 Artificial insemination.

Figure 9.2 Kidding percentage through AI at ICAR‐CIRG in different years.

Figure 9.3 Kids born through AI.

Chapter 11

Figure 11.1 Nondescript goat with triple kids.

Chapter 12

Figure 12.1 Stages of lactation.

Figure 12.2 Anatomical position, number, and teat morphology of goat.

Figure 12.3 Cistern of doe.

Figure 12.4 Physiology of milk ejection in goat.

Chapter 13

Figure 13.1 Reproductive system of a doe.

Figure 13.2 CIDR and its applicator.

Figure 13.3 Lutalyse

®

: a dinoprost trimethamine preparation (PGF2‐alpha...

Figure 13.4 GnRH preparation (Cystorelin

®

).

Figure 13.5 Somatic cell nuclear transfer in goats.

Chapter 14

Figure 14.1 Anestrus during the natural breeding season. Persistence of the ...

Figure 14.2 Possible causes leading to a failure of follicle development in ...

Figure 14.3 Hormonal patterns, types of ovarian cysts, and behavioral manife...

Figure 14.4 Causes of fertilization failure.

Figure 14.5 Pregnancy wastage in farm animals.

Figure 14.6 Causes of embryonic mortality.

Chapter 15

Figure 15.1 The uterus of a doe.

Figure 15.2 Structure of the ovary.

Chapter 17

Figure 17.1 Anatomical representation of a buck's reproductive system.

Chapter 18

Figure 18.1 Lymphatic system.

Chapter 19

Figure 19.1 Anatomical representation of a buck's reproductive system.

Chapter 20

Figure 20.1 Anatomy of the scrotum of small ruminants.

Chapter 21

Figure 21.1 Relationship of the seminiferous tubules to each other and to th...

Figure 21.2 Testes showing seminiferous tubules and excurrent ducts.

Figure 21.3 Sperm granuloma (epididymis exhibiting multiple sperm granulomas...

Figure 21.4 Irregular echogenicity indicates testicular pathology such as or...

Chapter 22

Figure 22.1 Implications of caprine abortion. ROP, retention of placenta.

Figure 22.2 Different malformations caused by Mimosa tenuiflora: (a) goat wi...

Figure 22.3 Hydrometra and true pregnancy.

Figure 22.4 Infectious causes of abortion in caprines.

Figure 22.5 Pathogenic mechanisms for induction of abortion in goats.

Chapter 23

Figure 23.1 Global seroprevalence rate of C. abortus (Tourin et al., 2022)....

Figure 23.2 Global seroprevalence of C. abortus in different animal species ...

Figure 23.3 Schematic diagram of Chlamydial developmental cycle. EB‐Elementa...

Figure 23.4 Different modes of transmission of C. abortus from infected shee...

Figure 23.5 Abortion in C. abortus infection is the combined effect of infla...

Figure 23.6 A. Almost full time aborted foetus with C. abortus infected doe....

Figure 23.7 Schematics diagram showing zoonotic transmission modes of C. abo...

Chapter 24

Figure 24.1 Landmarks in the discovery of Toxoplasma gondii.

Figure 24.2 Difficulties in estimating losses due to toxoplasmosis.

Figure 24.3 Routes of transmission.

Figure 24.4 The pathways of transmission of T. gondii are (a) cats (definiti...

Figure 24.5 The three infective stages of T. gondii are tachyzoite (a), brad...

Figure 24.6 (a) Longitudinal section view of the tachyzoite form of Toxoplas...

Figure 24.7 (a) Longitudinal section view of the bradyzoite form of Toxoplas...

Figure 24.8 (a) Longitudinal section view of the sporozoite form of Toxoplas...

Figure 24.9 (a) Tissue cyst of Toxoplasma gondii (3D animation), with a thic...

Figure 24.10 (a) Acquired ocular toxoplasmosis characterized by retinochoroi...

Chapter 25

Figure 25.1 Spread of Q fever among the population. 1. Aerosol. 2. Human con...

Figure 25.2 Transmission of Q fever (humans–animals–aerosol).

Figure 25.3 Different routes of infection in goats.

Figure 25.4 Pathogenesis of Q fever. 1. C. burnetii organisms. 2. Nascent Ch...

Figure 25.5 Clinical signs in goats with Q fever.

Figure 25.6 Goat with Q fever showing the aborted kid with changes in the pl...

Chapter 26

Figure 26.1 The various zoonotic Brucella species affecting humans.

Figure 26.2 The various clinical signs in goats with Brucella infection.

Figure 26.3 Gram‐stained Brucella under light microscopy.

Figure 26.4 Brucella species colony characteristics showing pinpoint, smooth...

Figure 26.5 RBPT testing of sera from animals suspected of brucellosis showi...

Figure 26.6 Various control strategies for brucellosis.

Chapter 27

Figure 27.1 Contamination by Campylobacter spp.

Chapter 28

Figure 28.1 Basic factors for noninfectious abortion in small ruminants.

Figure 28.2 Aspidosperma pyrifolium tree.

Figure 28.3 Ateleia glazioveana tree.

Figure 28.4 Tetrapterys multiglandulosa.

Figure 28.5 Enterolobium timbouva.

Figure 28.6 Stryphnodendron obovatum.

Figure 28.7 Stryphnodendron fissuratum. (a) Tree. (b) Leaves. (c) Seeds.

Figure 28.8 Mimosa tenuiflora.

Chapter 29

Figure 29.1 Causes of abortion in goats.

Figure 29.2 Aborted fetus of goat.

Chapter 30

Figure 30.1 Process of development of ketosis due to lack of feed intake.

Figure 30.2 (a–d) Lateral deviation of the neck in a pregnant doe.

Figure 30.3 Procedure of Rothera's test for detection of ketone bodies in ur...

Figure 30.4 Results of Rothera's test.

Chapter 31

Figure 31.1 Failure to meet nutritional requirements during the transition p...

Figure 31.2 Reasons for hypocalcemia in the pre‐ and postpartum periods.

Figure 31.3 Hyperexcitability and restlessness.

Figure 31.4 Doe in lateral recumbency.

Chapter 32

Figure 32.1 Complete uterine prolapse.

Figure 32.2 Incomplete uterine prolapse.

Figure 32.3 Uterine prolapse with visible cotyledon and endometrium.

Figure 32.4 Antiseptic washing of prolapsed uterus.

Figure 32.5 Repositioning of prolapsed uterine mass.

Figure 32.6 Uterine prolapse through vaginal tear exposing the serosal layer...

Figure 32.7 Class II rectal prolapse in goat.

Figure 32.8 Goat with applied truss.

Figure 32.9 Improvised retainer suture.

Chapter 33

Figure 33.1 Goat with retained fetal membrane.

Chapter 34

Figure 34.1 (a) Abnormal vaginal discharge in goat. (b) RFM in goat.

Chapter 35

Figure 35.1 Schematic representation of the development of postpartum uterin...

Figure 35.2 Risk factors for postpartum uterine infection.

Figure 35.3 Vaginoscope for use in sheep and goats.

Chapter 36

Figure 36.1 Mastitis in a goat.

Figure 36.2 (a) Inflamed udder. (b) Intramammary infusion. (c) Examination o...

Chapter 37

Figure 37.1 Individual kidding pen.

Figure 37.2 Clean straw bedding for a kid.

Figure 37.3 Goat kid enclosure.

Figure 37.4 FMD vaccine.

Figure 37.5 Goat pox vaccine.

Figure 37.6 Anthrax vaccine.

Figure 37.7 PPR vaccine.

Chapter 38

Figure 38.1 Sixty milliliter syringe and feeding tube.

Figure 38.2 Measurements before insertion.

Figure 38.3 Bottle feeding in goat kids.

Chapter 40

Figure 40.1 Heat homeostasis in goat kids.

Figure 40.2 Measurement of rectal temperature in a kid.

Figure 40.3 Management of hypothermia in the kid by fluid therapy.

Figure 40.4 Administration of milk replacer.

Chapter 41

Figure 41.1 Schematic representation of effects of exotoxins and endotoxins....

Chapter 42

Figure 42.1 Nasal discharge in kids.

Chapter 43

Figure 43.1 Schematic representation of scours caused by infectious etiology...

Figure 43.2 Schematic representation of scours caused by noninfectious etiol...

Figure 43.3 General transmission and pathogenesis of infectious pathogens ca...

Chapter 44

Figure 44.1 Schematic representation of the predisposing factors of joint il...

Chapter 45

Figure 45.1 Life cycle of goat coccidiosis.

Figure 45.2 Transmission pattern of coccidian occysts in goats.

Figure 45.3 Contributory factors in the development of coccidiosis.

Figure 45.4 (a) Pale congested mucous membrane. (b) Poor body weight in a go...

Figure 45.5 White nodules in intestinal mucosa in a goat affected with cocci...

Figure 45.6 Consequences of coccidiosis.

Figure 45.7 Unsporulated oocyst of Eimeria spp. in a goat.

Figure 45.8 An integrated approach to the management of coccidiosis.

Chapter 46

Figure 46.1 Difficulty in standing.

Figure 46.2 Reluctance to move or stand.

Figure 46.3 Muscle stiffness.

Figure 46.4 Poor growth in young animals.

Figure 46.5 Feeding a balanced diet.

Figure 46.6 Supplementation of selenium and/or vitamin E.

Chapter 47

Figure 47.1 Simplified diagrammatic representation of the pathogenesis of en...

Figure 47.2 A four‐month‐old goat kid affected by enterotoxemia revealing se...

Chapter 48

Figure 48.1 Etiology of acidosis in the goat.

Figure 48.2 Pathogenesis of acidosis.

Figure 48.3 Kid with diarrhea.

Figure 48.4 Death of kid due to severe acidosis.

Figure 48.5 Therapeutic management of acidosis by fluid administration in ki...

Chapter 49

Figure 49.1 Penis showing necrosis of the urethral process and glans due to ...

Figure 49.2 Urethra obstructed by uroliths at the sigmoid flexure showing ne...

Figure 49.3 Abundant uroliths and hemorrhagic cystitis with necrotic foci in...

Figure 49.4 Kidney presenting uroliths in the renal pelvis, hydronephrosis, ...

Chapter 50

Figure 50.1 Goat that died due to polioencephalomalacia showing typical posi...

Figure 50.2 Collection of cerebrospinal fluid from the cerebellomedullary ci...

Chapter 51

Figure 51.1 Diversity and source of transmission routes of Salmonella.

Figure 51.2 Black‐centered colonies of Salmonella spp. on bismuth sulfite ag...

Figure 51.3 Small, round, smooth‐edged, pink colonies of Salmonella spp. on ...

Chapter 52

Figure 52.1 Rectal prolapse in a goat.

Figure 52.2 Retention of prolapsed mass with purse‐string sutures.

Chapter 53

Figure 53.1 Common causes of copper deficiency.

Figure 53.2 Diagnosis of copper deficiency.

Figure 53.3 Treatment of copper deficiency.

Guide

Cover Page

Table of Contents

Title Page

Copyright Page

List of Contributors

Preface

Acknowledgment

Begin Reading

Index

WILEY END USER LICENSE AGREEMENT

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Elements of Reproduction and Reproductive Diseases of Goats

Edited by

Tanmoy Rana, MVSc, PhD

Assistant Professor, Department of Veterinary Clinical ComplexWest Bengal University of Animal & Fishery SciencesKolkata, India

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Library of Congress Cataloging‐in‐Publication Data Applied forHardback ISBN: 9781394189731ePDF: 9781394190072epub: 9781394190003oBook: 9781394190089

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List of Contributors

Mojetoluwa S. AfolabiDepartment of Food and Animal ScienceAlabama A&M University, Normal, USA

Akanksha AgnihotriDepartment of Veterinary MedicineCollege of Veterinary Science and Animal HusbandryJabalpur, India

Manoj Kumar AhirwarDepartment of Veterinary Surgery and RadiologyCollege of Veterinary Science and Animal HusbandryJabalpur, India

Allam LushaikyaaFaculty of Veterinary MedicineAhmadu Bello UniversityZaria, Nigeria

Varun AsediyaAnimal Nutrition Research StationAnand Agricultural UniversityAnand, Gujarat, India

Ashok AshwiniDepartment of Veterinary MedicineVeterinary CollegeHassan, India

Ashok BaindhaPost Graduate Institute of VeterinaryEducation & ResearchJaipur, India

K. AthiraDepartment of Veterinary Epidemiologyand Preventive MedicineCollege of Veterinary and Animal SciencesMannuthy, India

Kolawole Jonathan BamideleFaculty of Veterinary MedicineAhmadu Bello UniversityZaria, Nigeria

Himangshu BaruahDepartment of Veterinary Pharmacology and ToxicologyLakhimpur College of Veterinary ScienceAssam Agricultural UniversityNorth Lakhimpur, India

Sonam BhattDepartment of Veterinary MedicineBihar Veterinary College, Bihar Animal Sciences UniversityPatna, India

Suman BiswasTeaching Veterinary Clinical ComplexFaculty of Veterinary and Animal SciencesInstitute of Agricultural ScienceBanaras Hindu UniversityMirzapur, India

Antonio Carlos Lopes CâmaraLarge Animal Veterinary Teaching HospitalUniversidade de BrasíliaBrasília, Brazil

Vipin ChandraPost Graduate Institute of VeterinaryEducation & ResearchJaipur, India

Prakash M. ChauhanDepartment of Clinics, College of Veterinary Scienceand Animal HusbandryKamdhenu University, SardarkrushinagarGujarat, India

Akanksha ChoudharyPost Graduate Institute of VeterinaryEducation & ResearchJaipur, India

Bashir Maryam DarmaVeterinary Teaching HospitalFaculty of Veterinary MedicineBayero UniversityKano, Nigeria

Bhupamani DasDepartment of ClinicsCollege of Veterinary Science and Animal HusbandryKamdhenu UniversitySardarkrushinagar, Gujarat, India

Jitumoni DasDepartment of Animal ReproductionGynaecology and ObstetricsLakhimpur College of Veterinary ScienceAssam Agricultural UniversityNorth Lakhimpur, India

K. Justin DavisDepartment of Veterinary Epidemiologyand Preventive MedicineCollege of Veterinary and Animal SciencesMannuthy, India

Sarita DeviDepartment of MedicineCollege of Veterinary Science and Animal HusbandryKamdhenu UniversitySardarkrushinagar, Gujarat, India

Kritika DhialApollo College of Veterinary MedicineJaipur, Rajasthan, India

Priyanka DuggalDepartment of Veterinary Surgery and RadiologyCollege of Veterinary SciencesLala Lajpat Rai University of Veterinaryand Animal SciencesHisar, India

Parimita DuttaKrishi Vigyan Kendra (KVK)Tirap, India

Ayyasamy ElangoDepartment of Veterinary Gynaecology and ObstetricsVeterinary College and Research InstituteTamil Nadu Veterinary and Animal Sciences UniversitySalem, India

Ama Adadzewa EshunDepartment of Food and Animal ScienceAlabama A&M University, Normal, USA

Abhijeet FernandesDepartment of Veterinary Gynaecology and ObstetricsCollege of Veterinary ScienceGuru Angad Dev Veterinary and Animal Sciences UniversityBathinda, India

Chetana GangwarAnimal Physiology and Reproduction DivisionICAR‐Central Institute for Research on GoatMathura, India

Amaan GhaaziApollo College of Veterinary MedicineJaipur, Rajasthan, India

Mohammad M. IslamPashupalan Sanshodhan KendraKamdhenu UniversityRamna Muvada, Gujarat, India

Manoj Kumar KalitaDepartment of Animal ReproductionGynaecology and ObstetricsLakhimpur College of Veterinary ScienceAssam Agricultural UniversityNorth Lakhimpur, India

Laishram Sanathoi KhumanDepartment of Extension EducationLakhimpur College of Veterinary ScienceAssam Agricultural UniversityNorth Lakhimpur, India

Hycinth Ndabatsado KoloDepartment of Animal Health and ProductionFederal University of TechnologyMinna, Nigeria

Anil KumarDepartment of Veterinary Clinical ComplexBihar Veterinary College, Bihar Animal Sciences UniversityPatna, India

Ankesh KumarDepartment of Veterinary Clinical ComplexBihar Veterinary College, Bihar Animal Sciences UniversityPatna, India

Gowdaiahnadoddi K. Chetan KumarDepartment of Veterinary MedicineVeterinary CollegeHassan, India

Kaushalendra KumarDepartment of Animal NutritionBihar Veterinary CollegePatna, India

Pramod KumarDepartment of Veterinary Physiology and BiochemistryCollege of Veterinary Science and Animal HusbandryAcharya Narendra Deva University ofAgriculture and TechnologyAyodhya, Uttar Pradesh, India

Rajesh KumarDepartment of Veterinary Gynaecology and ObstetricsCollege of Veterinary Science and Animal HusbandryAcharya Narendra Deva University ofAgriculture and TechnologyAyodhya, India

Rohit KumarDepartment of Livestock Production ManagementDr. G.C. Negi College of Veterinary and Animal SciencesPalampur, India

Sanjay KumarDepartment of Animal NutritionBihar Veterinary CollegePatna, India

Rajni KumariICAR‐RCER, Patna, India

Neeti LakhaniDepartment of Animal NutritionCollege of Veterinary ScienceGuru Angad Dev Veterinary and Animal Sciences UniversityBathinda, India

Mohan LalPost Graduate Institute of VeterinaryEducation & ResearchJaipur, India

Mada A. KhumranFaculty of Veterinary MedicineAhmadu Bello UniversityZaria, Nigeria

Chinmoy MajiDivision of Animal Health, North 24 Parganas KrishiVigyan Kendra, West Bengal University of Animal &Fishery Sciences, Ashokenagar, West Bengal, India

Kruti Debnath MandalDepartment of Avian SciencesFaculty of Veterinary & Animal SciencesWest Bengal University of Animal & Fishery SciencesNadia, India

Sakthivel ManokaranDepartment of Veterinary Gynaecology and ObstetricsVeterinary College and Research Institute, Tamil NaduVeterinary and Animal Sciences UniversitySalem, India

Varsha MishraDepartment of Veterinary MedicineCollege of Veterinary Science and Animal HusbandryJabalpur, India

Aliyu Abdullahi MohammedDepartment of Animal SciencesUsman Danfodio UniversitySokoto, Nigeria

Ogunkunle NathanielDepartment of Food and Animal ScienceAlabama A&M UniversityNormal, USA

Richard Jagatheesan Pitchai NavlonVeterinary College and Research InstituteTamil Nadu Veterinary and Animal Sciences UniversityTheni, India

Pauline Chiezey NgoziNational Animal Production Research Institute/Faculty of Veterinary Medicine, Ahmadu Bello UniversityZaria, Nigeria

Ram NiwasDepartment of Veterinary Surgery and RadiologyCollege of Veterinary SciencesLala Lajpat Rai University of Veterinary and Animal SciencesHisar, India

Adeniyi NjidekaDepartment of Biological Sciences, Alabama A&MUniversity, Normal, USA

Padmanath KrishnanVeterinary Clinical ComplexVeterinary College and Research InstituteTamil Nadu Veterinary and Animal Sciences UniversityTheni, India

Chirag P. ParmarPashupalan Sanshodhan KendraKamdhenu UniversityRamna Muvada, Gujarat, India

Husen R. ParsaniDepartment of Veterinary ParasitologyCollege of Veterinary Science and Animal HusbandryKamdhenu UniversitySardarkrushinagar, Gujarat, India

Kashyap PatelDepartment of MedicineCollege of Veterinary Science and Animal HusbandryKamdhenu UniversitySardarkrushinagar, Gujarat, India

Ramesh M. PatelDepartment of ClinicsCollege of Veterinary Science and Animal HealthKamdhenu UniversitySardarkrushinagar, Gujarat, India

Abhishek PathakApollo College of Veterinary MedicineJaipur, Rajasthan, India

Salil Kumar PathakDepartment of Veterinary MedicineCollege of Veterinary Science and Animal HusbandryJabalpur, India

Prerona PatowaryDepartment of Veterinary Clinical Medicine, Ethics andJurisprudence, Assam Agricultural University – GoatResearch Station, Burnihat, India

Shashi PradhanDepartment of Veterinary MedicineCollege of Veterinary Science and Animal HusbandryJabalpur, India

Ankit S. PrajapatiVeterinary Clinical Complex (Medicine), College ofVeterinary Science and Animal Health, KamdhenuUniversity, Sardarkrushinagar, Gujarat, India

Ravi D. PrajapatiDepartment of Veterinary Gynaecology and ObstetricsCollege of Veterinary Science and Animal HusbandryKamdhenu UniversitySardarkrushinagar, India

Aditya PratapDepartment of Veterinary MedicineCollege of Veterinary Science and Animal HusbandryJabalpur, India

Saahithya RajamohanDepartment of Veterinary PathologyVeterinary College and Research InstituteTamil Nadu Veterinary and Animal Sciences UniversitySalem, India

Ranjani RajasekaranDepartment of Veterinary MicrobiologyVeterinary College and Research InstituteTamil Nadu Veterinary and Animal Sciences UniversityTheni, India

Nirmal Singh RajawatCollege of Veterinary and Animal SciencesRajasthan University of Veterinary and Animal SciencesBikaner, India

Ramasamy RajkumarDepartment of Veterinary Gynaecology and ObstetricsVeterinary College and Research InstituteTamil Nadu Veterinary and Animal Sciences UniversitySalem, India

Jeetendra S. RajoriyaDepartment of Veterinary Gynaecology and ObstetricsCollege of Veterinary Science and Animal HusbandryJabalpur, India

Tanmoy RanaDepartment of Veterinary Clinical ComplexWest Bengal University of Animal and Fishery SciencesKolkata, India

Ravi RanjanAnimal Physiology and Reproduction DivisionICAR‐Central Institute for Research on GoatMathura, India

Jigar RavalNational Dairy Development BoardAnand, India

Sunant RavalCollege of Veterinary Science and Animal HusbandryKamdhenu UniversityAnand, India

Meesam RazaDepartment of Livestock Production ManagementDr. G.C. Negi College of Veterinary and Animal SciencesPalampur, India

Bhavanam Sudhakara ReddyCollege of Veterinary Science, Sri VenkateswaraVeterinary University, Proddatur, Andhra Pradesh, India

Peter Ibrahim RekwotFaculty of Veterinary MedicineAhmadu Bello UniversityZaria, Nigeria

Abdul ReshmaDepartment of Veterinary Gynaecology and ObstetricsVeterinary College and Research InstituteTamil Nadu Veterinary and Animal Sciences UniversitySalem, India

Amitava RoyDepartment of Livestock Farm ComplexWest Bengal University of Animal & Fishery SciencesKolkata, India

Vikas SachanDepartment of Veterinary Gynaecology, DUVASUMathura, India

Rayala S. SagarDepartment of Veterinary MedicineVeterinary CollegeHassan, India

Gitesh SainiDepartment of Veterinary Gynaecology and ObstetricsCollege of Veterinary ScienceGuru Angad Dev Veterinary and Animal Sciences UniversityBathinda, India

Felix Uchenna SamuelFaculty of Veterinary MedicineNational Animal Production Research InstituteAhmadu Bello UniversityZaria, Nigeria, and

Department of Food and Animal ScienceAlabama A&M UniversityNormal, USA

Tulasiraman SarathDepartment of Veterinary Gynaecology and ObstetricsVeterinary College and Research InstituteTamil Nadu Veterinary and Animal Sciences UniversitySalem, India

Pardeep SharmaDepartment of Veterinary MedicineDr G.C. Negi College of Veterinary and Animal SciencesPalampur, India

Idris SherifFaculty of Veterinary MedicineAhmadu Bello UniversityZaria, Nigeria

Abhiyuday P. SinghDepartment of Veterinary Gynaecology & ObstetricsCollege of Veterinary Science and Animal HusbandryJabalpur, India

Brejesh SinghDepartment of Veterinary MedicineCollege of Veterinary Science and Animal HusbandryJabalpur, India

Rashmi SinghPost Graduate Institute of VeterinaryEducation & ResearchJaipur, India

Satyavrat V. SinghDepartment of Veterinary MedicineCollege of Veterinary Science and Animal HusbandryAcharya Narendra Deva University ofAgriculture and TechnologyAyodhya, Uttar Pradesh, India

Bhoomika SirsantDepartment of Veterinary Public Health andEpidemiology, Bihar Veterinary CollegeBihar Animal Sciences UniversityPatna, India

Sirigireddy SivajothiCollege of Veterinary Science, Sri VenkateswaraVeterinary UniversityProddatur, Andhra Pradesh, India

Simson SorenDepartment of Veterinary PhysiologyLakhimpur College of Veterinary ScienceAssam Agricultural UniversityNorth Lakhimpur, India

Benito Soto‐BlancoDepartment of Veterinary Clinics and SurgeryVeterinary SchoolUniversidade Federal de Minas GeraisBelo Horizonte, Brazil

Abhinav N. SutharDepartment of MedicineCollege of Veterinary Science and Animal HusbandryKamdhenu UniversitySardarkrushinagar, Gujarat, India

Ankaj ThakurDepartment of Livestock Production ManagementDr G.C. Negi College of Veterinary and Animal SciencesPalampur, India

Annadurai ThangamaniDepartment of Veterinary Gynaecology and ObstetricsVeterinary College and Research InstituteTamil Nadu Veterinary and Animal Sciences UniversitySalem, India

Amita TiwariDepartment of Veterinary MedicineCollege of Veterinary Science and Animal HusbandryJabalpur, India

Mahendra P.S. TomarDepartment of Veterinary AnatomyNTR College of Veterinary ScienceGannavaram, India

Shivangi UdainiyaVeterinary Polytechnic CollegeNanaji Deshmukh Veterinary Science UniversityBhopal, India, andDepartment of Veterinary MedicineCollege of Veterinary Science and Animal HusbandryJabalpur, India

Hridya Susan VarugheseDepartment of Veterinary MicrobiologyVeterinary College, Karnataka Veterinary Animal andFisheries Sciences UniversityHebbal, India

Abhinov VermaDepartment of Veterinary Anatomy, DUVASUMathura, India

Prerna YadavPost Graduate Institute of VeterinaryEducation and ResearchJaipur, India

Somu YogeshpriyaDepartment of Veterinary MedicineVeterinary College and Research InstituteChennai, India

Preface

This book provides an essential resource for students, academicians, and practitioners and will be beneficial for audiences in the farm and industry sectors, and milk producers' union. The book provides practical, updated information about the normal reproduction, reproductive diseases, and management of goats. It covers the current thoughts and scientific modules for the betterment of understanding of the readers. The book illustrates the reproductive anatomy and physiology of both males and females in a systematic way.

The book is divided into three parts: normal reproduction, reproductive disorders, and reproductive management of goats. The main topics covered include breeding characteristics, pregnancy detection, diagnosis, treatment, and management of infertility, obstetrics, abortion, and care and management of kids. The book emphasizes current knowledge about the physiological and biochemical mechanisms in the regulation of reproduction. It will serve as a reference book for students, veterinary nurses, researchers, academicians, scientists, and professional workers engaged in small ruminant production, applicable to anyone who has a keen interest in the biology of normal reproduction, and reproductive diseases, and management of goats. Artificial insemination and advanced reproductive biotechnology are also described and discussed. Featuring contributions from experts in various fields, each section covers clinically relevant aspects of the reproductive anatomy and physiology of both males and females.

The main important topics include the evaluation of the normal physiological function of reproductive organs, breeding soundness, pregnancy diagnosis, diagnosis and treatment of infertility, abortion, obstetrics, surgery of the reproductive tract, care of kids, and the latest reproductive technology, illustrated throughout with figures and tables.

Tanmoy Rana

Department of Veterinary Clinical Complex, West BengalUniversity of Animal & Fishery Sciences, Kolkata, India

Acknowledgment

I wish to thank all the authors of the first edition for their hard work, and sharing their expertise and knowledge to make the book fruitfully. I also express my gratitude to our Honourable Vice Chancellor, West Bengal University of Animal & Fishery Sciences, Kolkata, India, for giving me the opportunity to edit the book. I also convey my sincere gratitude and wholehearted thanks to all veterinarians, farm managers, and academicians for their expert opinions and thoughts to make the book a success. I would like to express my sincere regards to Dr Rituparna Bose, Commissioning Editor, Bhavya Boopathi, Managing Editor, and other members of Wiley Publishers who actively helped me during the editing of the book. I would also like to thank my family members for giving me great support and time to finalize the book.

Tanmoy Rana

Kolkata, India

Section 1Goat Reproductive Health

1Anatomy of the Reproductive System in the Buck and Doe

Abhinov Verma1, Mahendra P.S. Tomar2, and Vikas Sachan3

1 Department of Veterinary Anatomy, DUVASU, Mathura, India

2 Department of Veterinary Anatomy, NTR College of Veterinary Science, Gannavaram, India

3 Department of Veterinary Gynaecology, DUVASU, Mathura, India

1.1 Introduction

Reproduction is crucial for the survival of species. It is the method used by living things to reproduce (Frandson et al. 2003). Since being domesticated, the goat has become seasonally polyestrous, meaning that it reproduces naturally at specific periods of the year. The photoperiod, rasse, and nutrition are the key determinants of this trait's variation (Delgadillo et al. 2012). Several areas of goat production are influenced by reproduction, so managing reproduction requires an understanding of this process. For survival of the species, the production of meat, milk, skin, and fiber, as well as the replenishment of breeding stock, a high rate of reproductive efficiency is crucial (Abebe 2008).

Male and female individuals in each species have specific anatomical and functional features (Nickel et al. 1973). Therefore, for easier understanding, this chapter is divided into two sections: the male reproductive system and female reproductive system in goats.

1.2 Male Reproductive System

The major function of this system is the production of spermatozoa and male sex hormone (Frandson et al. 2003). The male reproductive system consists of the following structures (Figures 1.1 and 1.2).

Scrotum

Testes

Epididymis

Ductus deferens

Accessory sex glands

Penis

Urethra

1.2.1 Scrotum

The testicles and nearby spermatic cord segments are located in the scrotum, which is a sac or pouch (Figure 1.1). It can take on a variety of shapes depending on the health of its subcutaneous muscular tissue. When exposed to cold, the latter constricts, causing the scrotum to rise and become thicker and wrinkled; when relaxed by heat, weariness, or debility, however, it becomes smooth and pendulous with a superior neck constriction (Frandson et al. 2003). It consists of layers which correspond with those of the abdominal wall, considered from without inward: skin, dartos, scrotal fascia, parietal layer of the tunica vaginalis (Getty 1975; Farooqui 2004; Pathak et al. 2014).

Figure 1.1 Diagrammatic presentation of the male reproductive system.

Figure 1.2 Photograph showing the organs of the male reproductive system.

The spermatic cord begins at the abdominal inguinal ring, where its constituent parts come together, extends obliquely downward through the inguinal canal, passes over the side of the penis and ends at the attached border of the testicle (Figure 1.2). It consists of the following structures.

Spermatic artery.

Spermatic vein.

Lymphatics.

Sympathetic nerves.

Ductus deferens.

Internal cremaster muscles.

Visceral layer of tunica vaginalis.

1.2.2 Testes

The testis is the seat of production of male germ cells. The testes are located in the inguinal region of the body and are enclosed in the scrotum (Frandson et al. 2003; Pathak 2006). They descend into the scrotum from the abdominal cavity during fetal development. Descent of the testes from the genital ridge in the fetus to an extracorporeal location after birth is a mandatory development process to ensure normal spermatogenesis in the mature testis (Farooqui 2004).

Each testis weighs about 130–160 g with an elongated oval form and the long axis nearly vertical (Figure 1.1). It comprises two surfaces, two borders and two extremities. Both the surfaces, i.e. the medial and lateral surfaces, are convex and smooth. The cranial border is free and the caudal border is attached to the epididymis. The dorsal and ventral extremities are termed the head and tail, respectively. A testicle of an average sized adult buck is 6–7, 4–5, and 4–5 cm in length, width, and thickness (in middle), respectively (Solaiman 2010). The testes are frequently of unequal size, with the left typically being larger, and their sizes fluctuate greatly. Testicular growth is extremely rapid from birth to puberty and then does not change significantly (Pathak et al. 2014). The testes of goats are broader in proportion to their length.

Figure 1.3 Photograph showing the testis and tunica vaginalis.

The tunica vaginalis, which is the visceral layer of the serous envelope surrounding the spermatic cord and testicle, covers the majority of the testicle's surface (Figure 1.3). This is reflected from the attached border of the gland, leaving an exposed area where the spermatic cord's vessels and nerves enter the testicle. The tunica albuginea, a robust capsule made of dense white fibrous tissue and unstriped muscle fibers, lies beneath the tunica vaginalis.

The spermatic artery supplies blood to the testicle. The artery is quite convoluted close to the testicle and descends in the anterior portion of the spermatic cord. It splits into many branches that enter the gland on the trabeculae and septa (septula testes) and ascend and descend on the testicular surfaces (Figure 1.4). The pampiniform plexus is a network formed by the testicular veins as they exit the testis and encircle the spermatic cord's artery. The lymphatic vessels often follow the veins' path before entering the lumbar lymph glands. The spermatic plexus, to which the vessels are primarily dispersed, is formed by the renal and posterior mesenteric plexuses, which supply the nerves.

The exocrine function of the testes is spermatogenesis under the influence of follicle‐stimulating hormone (FSH) and testosterone, whereas their endocrine function is the synthesis and release of testosterone by Leydig cells under the effect of luteinizing hormone (LH)/interstitial cell‐stimulating hormone (ICSH). Castration is removal of the testicles in order to reduce buck aggression (König and Liebich 2020).

Figure 1.4 Photograph (sagittal section) showing the septula testis.

The blood supply to the testes is via the internal spermatic artery and innervation occurs via the renal and mesenteric plexuses.

1.2.3 Epididymis

The epididymis serves as the chief ductal system, besides providing nutrition for growth and maturation. The epididymis is attached to the caudal border of the testicle and somewhat overlaps the lateral surface. Its dorsal enlarged end is termed the head (caput epididymis), its ventral, slightly enlarged end is the tail (cauda epididymis), and the intermediate part is the body (corpus epididymis). The head is closely connected with the testicle by the efferent ducts, in continuation of the rete testis. The efferent ducts together form the ductus epididymis which runs inside the body of the epididymis. The ductus epididymis is very flexible and almost 60 m in length in the adult buck. The tail is continued by the ductus deferens/vas deferens which is attached to the posterior extremity of the testicle by the ligamentum epididymis.

1.2.4 Ductus Deferens

The ductus deferens extends from the tail of the epididymis to the pelvic urethra. It ascends in the inguinal canal, enclosed within the spermatic cord. At the vaginal ring, it separates from the other constituents of the spermatic cord, and turns backward and inward into the pelvic cavity to reach the pelvic urethra just caudal to the neck of the bladder. The two ducts lie very close together and empty on the roof of the urethra as enlarged ampullae.

1.2.5 Accessory Sex Glands or Annex Glands

Accessory sex glands are conspicuous outgrowths of the genital tract and can be regarded as part of the male reproductive system as they play an integral role in the fertility process. In the buck, they include the vesicular gland, prostate gland and bulbourethral or Cowper's gland.