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A practical and up-to-date guide to canine reproduction for dog breeders, veterinary students and veterinary surgeons. Written in an easy-to-use style and presented in a practically useful format with clear illustrations. The information is supported by references from the author’s published work and his wealth of clinical experience to form the important authoritative text in this field. A vital aid to those seeking an understanding of normal and abnormal reproduction as well as common reproductive techniques.
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Seitenzahl: 490
Veröffentlichungsjahr: 2012
Table of Contents
Cover
Title page
Copyright page
Preface
Section 1: Breeding, Reproduction and Care of the Normal Animal
1 Care of the Juvenile, Genetics and Planning of Breeding
1.1 BEHAVIOURAL DEVELOPMENT
1.2 HEALTH CARE
1.3 PHYSIOLOGICAL DEVELOPMENT
1.4 ONSET OF PUBERTY
1.5 SELECTION OF ANIMALS FOR BREEDING
1.6 BASIC GENETICS
1.7 BREEDING SCHEMES
1.8 FIT TO BREED
1.9 BREEDING AT THE FIRST (PUBERTAL) OESTRUS
2 Anatomy of the Female
2.1 VULVA
2.2 VESTIBULE
2.3 VESTIBULO-VAGINAL JUNCTION
2.4 VAGINA
2.5 CERVIX UTERI
2.6 UTERUS
2.7 UTERINE (FALLOPIAN) TUBES
2.8 OVARIES
2.9 MESENTERY
2.10 BLOOD SUPPLY
2.11 MAMMARY GLAND
3 Physiology of the Female
3.1 TERMINOLOGY
3.2 PRE-PUBERTY
3.3 PRO-OESTRUS
3.4 OESTRUS
3.5 METOESTRUS (DIOESTRUS)
3.6 ANOESTRUS
3.7 FALSE PREGNANCY
4 Endocrinology of the Female
4.1 GONADOTROPHIN-RELEASING HORMONE (GnRH, RARELY TERMED LHRH)
4.2 FOLLICLE-STIMULATING HORMONE (FSH)
4.3 LUTEINIZING HORMONE (LH)
4.4 OESTROGENS
4.5 PROGESTERONE
4.6 PROLACTIN
4.7 ANDROGENS
4.8 PROSTAGLANDINS
5 Clinical Examination of the Female
5.1 BASIC EXAMINATION OF THE REPRODUCTIVE TRACT
5.2 ADDITIONAL TECHNIQUES THAT MAY BE CONSIDERED
5.3 VISUAL EXAMINATION OF THE PERINEUM
5.4 PALPATION OF THE MAMMARY GLANDS
5.5 VAGINAL EXAMINATION
5.6 DIGITAL EXAMINATION OF THE REPRODUCTIVE TRACT
5.7 ENDOSCOPIC EXAMINATION OF THE FEMALE REPRODUCTIVE TRACT
5.8 ULTRASONOGRAPHIC EXAMINATION OF THE FEMALE REPRODUCTIVE TRACT
5.9 MEASUREMENT OF PLASMA HORMONES
5.10 BACTERIAL EXAMINATION OF THE FEMALE REPRODUCTIVE TRACT
5.11 CYTOLOGICAL EXAMINATION OF SUSPECTED PATHOLOGICAL MATERIAL
5.12 ENDOCRINOLOGICAL STIMULATION TESTS
5.13 LAPAROSCOPY AND LAPAROTOMY
5.14 KARYOTYPING
6 Anatomy of the Male
6.1 PREPUCE (SHEATH)
6.2 PENIS
6.3 TESTES
6.4 EPIDIDYMIS
6.5 DUCTUS DEFERENS (VAS DEFERENS)
6.6 SPERMATIC CORD
6.7 INGUINAL CANAL
6.8 PROSTATE GLAND
6.9 URETHRA
7 Physiology of the Male
7.1 SPERMATOGENESIS
7.2 TEMPERATURE CONTROL
7.3 SPERMATOZOAL TRANSPORT
7.4 PROSTATE GLAND
7.5 ERECTION
7.6 EJACULATION
7.7 SPERM TRANSPORT IN THE BITCH
8 Endocrinology of the Male
8.1 GONADOTROPHIN-RELEASING HORMONE (GnRH)
8.2 FOLLICLE-STIMULATING HORMONE (FSH)
8.3 LUTEINIZING HORMONE (LH) (ALSO CALLED INTERSTITIAL CELL STIMULATING HORMONE (ICSH))
8.4 ANDROGENS (TESTOSTERONE AND DIHYDROTESTOSTERONE)
8.5 INHIBIN
8.6 ACTIVINS AND INHIBINS
9 Normal Mating
9.1 PLANNING THE MATING
9.2 TIMING
9.3 PREPARATION FOR MATING
9.4 NORMAL COITAL BEHAVIOUR
9.5 FACTORS PREVENTING NORMAL MATING
9.6 CERTIFICATE OF MATING
10 Clinical Examination of the Male
10.1 BASIC EXAMINATION OF THE REPRODUCTIVE TRACT
10.2 ADDITIONAL TECHNIQUES THAT MAY BE CONSIDERED
10.3 VISUAL EXAMINATION OF THE PENIS AND PREPUCE
10.4 PALPATION OF THE SCROTAL CONTENTS
10.5 SEMEN COLLECTION AND EVALUATION
10.6 MEASUREMENT OF ALKALINE PHOSPHATASE CONCENTRATIONS
10.7 ULTRASONOGRAPHIC EXAMINATION OF THE MALE REPRODUCTIVE TRACT
10.8 MEASUREMENT OF PLASMA HORMONES
10.9 OBSERVATION OF MATING BEHAVIOUR
10.10 BACTERIAL EXAMINATION OF THE MALE REPRODUCTIVE TRACT
10.11 TESTICULAR BIOPSY
10.12 TESTICULAR FINE-NEEDLE ASPIRATION
10.13 PROSTATIC BIOPSY
10.14 PROSTATIC MASSAGE/URETHRAL FLUSHING
10.15 ENDOCRINOLOGICAL STIMULATION TESTS
10.16 KARYOTYPING
11 Fertilization and the Optimal Breeding Time
11.1 MATING TIME AND FERTILIZATION TIME
11.2 REPRODUCTIVE PHYSIOLOGY
11.3 ASSESSING THE OPTIMAL TIME FOR MATING
12 Pregnancy and Lactation
12.1 PREGNANCY LENGTH
12.2 ENDOCRINE CHANGES IN BLOOD
12.3 DEVELOPMENT OF THE CONCEPTUS
12.4 DEVELOPMENT OF THE PLACENTA
12.5 OTHER CHANGES DURING PREGNANCY
12.6 PRODUCTION, MAINTENANCE AND EJECTION OF MILK
13 Pregnancy Diagnosis
13.1 ABSENCE OF A RETURN TO OESTRUS
13.2 BEHAVIOURAL CHANGES
13.3 PHYSICAL CHANGES
13.4 ABDOMINAL PALPATION
13.5 IDENTIFICATION OF FETAL HEART BEATS
13.6 RADIOGRAPHY
13.7 ENDOCRINOLOGICAL TESTS
13.8 ACUTE PHASE PROTEINS
13.9 ULTRASOUND EXAMINATION
14 Normal Whelping
14.1 ENDOCRINE CONTROL
14.2 PREPARATION OF THE ENVIRONMENT
14.3 EQUIPMENT
14.4 PREPARATION OF THE BITCH
14.5 THE ‘OVERDUE’ BITCH
14.6 FIRST STAGE PARTURITION
14.7 SECOND STAGE PARTURITION
14.8 THIRD STAGE PARTURITION
14.9 INDUCTION OF PARTURITION
14.10 CARE OF THE BITCH AFTER PARTURITION
15 Care of Neonatal Puppies
15.1 PHYSIOLOGY AND HUSBANDRY
15.2 BEHAVIOUR
15.3 APPEARANCE OF THE NORMAL HEALTHY NEONATAL PUP
15.4 CARE OF NEONATES AFTER PARTURITION
Section 2: Abnormalities of Reproduction and the Perinatal Period
16 Abnormalities of the Female
16.1 VULVA
16.2 VESTIBULE
16.3 VAGINA
16.4 CERVIX
16.5 UTERUS
16.6 UTERINE TUBES
16.7 OVARIES
16.8 MAMMARY GLAND
16.9 URETHRA
17 Abnormalities of the Male
17.1 PREPUCE (SHEATH)
17.2 PENIS
17.3 TESTES
17.4 EPIDIDYMIDES
17.5 VASA DEFERENTIA
17.6 URETHRA
17.7 PROSTATE GLAND
17.8 SCROTUM
17.9 MAMMARY GLAND
18 Intersex Animals
18.1 DEFINITIONS
18.2 AETIOLOGY
18.3 CLINICAL SIGNS
18.4 EXTERNAL PHYSICAL SIGNS
18.5 INTERNAL REPRODUCTIVE ORGANS
18.6 TREATMENT
19 Abnormal Female Physiology and Endocrinology
19.1 DELAYED PUBERTY (PRIMARY ANOESTRUS)
19.2 PROLONGED FIRST PRO-OESTRUS
19.3 PROLONGED PRO-OESTRUS OR OESTRUS
19.4 SHORT INTER-OESTROUS PERIODS (OVULATION FAILURE)
19.5 PROLONGED ANOESTRUS (SECONDARY ANOESTRUS)
19.6 UNPREDICTABLE OVULATION TIME
19.7 SILENT OESTRUS
19.8 PSEUDOPREGNANCY
19.9 THYROID DEFICIENCY
19.10 OTHER HORMONAL ABNORMALITIES
20 Abnormal Male Physiology and Endocrinology
20.1 HYPOGONADISM
20.2 POOR LIBIDO
20.3 ABSENCE OF SPERMATOGENESIS
20.4 OTHER CAUSES OF ABNORMAL SPERMATOGENESIS
20.5 IMPAIRED SPERMATOZOAL TRANSPORT
20.6 IMPOTENCE (INABILITY TO COPULATE)
20.7 ATTRACTIVENESS TO OTHER MALE DOGS
21 Abnormal Pregnancy
21.1 FERTILIZATION FAILURE
21.2 EARLY EMBRYONIC DEATH
21.3 RESORPTION
21.4 UTERINE BODY PREGNANCY
21.5 ABORTION
21.6 FETAL ABNORMALITIES
21.7 METROCOELE
21.8 UTERINE RUPTURE
21.9 LOSS OF FETAL FLUID
21.10 PREGNANCY DIABETES
21.11 HYPOGLYCAEMIA
22 Abnormal Parturition and Problems After Whelping
22.1 CLASSIFICATION OF DYSTOCIA
22.2 THE ‘OVERDUE’ BITCH
22.3 BLACK/GREEN DISCHARGE WITH NO SIGNS OF PARTURITION
22.4 NERVOUS INHIBITION OF PARTURITION
22.5 INEFFECTUAL STRAINING FOR OVER 1 HOUR
22.6 BITCH HAD SEVERAL PUPPIES BUT RESTLESS OR LARGER LITTER EXPECTED
22.7 OXYTOCIN TREATMENT
22.8 OBSTETRICAL MANIPULATION
22.9 UTERINE RUPTURE
22.10 UTERINE TORSION
22.11 RETAINED FETAL MEMBRANES
22.12 POST-PARTUM METRITIS
22.13 POST-PARTUM HAEMORRHAGE
22.14 SUBINVOLUTION OF PLACENTAL SITES
22.15 HYPOCALCAEMIA (PUERPERAL TETANY OR ECLAMPSIA)
22.16 CANNIBALISM
22.17 PROLAPSED UTERUS
22.18 FAILURE OF MILK LET-DOWN
22.19 AGALACTIA
22.20 MASTITIS
22.21 GALACTOSTASIS
23 Care of the Sick Neonate
23.1 IMPORTANT CONSIDERATIONS
23.2 SIGNS THAT THE NEONATE MAY BE SICK
23.3 TREATING A SICK PUP WITH MINOR SIGNS OF DISEASE
23.4 TREATING A SICK PUP WITH SERIOUS SIGNS OF DISEASE
23.5 CAUSES OF DEATH IN NEONATES
23.6 COMMON CAUSES OF ILLNESS IN NEONATES
24 Approach to Infertility Cases
24.1 DEFINITIONS
24.2 EXPECTATIONS OF FERTILITY
24.3 THE INFERTILE BITCH
24.4 TREATMENT OF INFERTILITY IN THE BITCH
24.5 THE INFERTILE DOG
24.6 TREATMENT OF INFERTILITY IN THE DOG
24.7 INFERTILITY IN KENNELS
24.8 WHY DO WE KNOW SO LITTLE ABOUT INFERTILITY IN DOGS?
Section 3: Control of Reproduction and Reproductive Technologies
25 Hormonal Contraception
25.1 ADVANTAGES OF PREVENTION AND SUPPRESSION OF OESTRUS
25.2 DISADVANTAGES OF PREVENTION AND SUPPRESSION OF OESTRUS
25.3 SUPPRESSION OF A PRO-OESTRUS THAT HAS ALREADY STARTED
25.4 PREVENTION OF AN EXPECTED PRO-OESTRUS/OESTRUS
25.5 PREVENTION OF REPRODUCTIVE BEHAVIOUR FOR EXTENDED PERIODS OF TIME
25.6 FUTURE METHODS OF PREVENTING OESTRUS
25.7 UNWANTED MATING
25.8 HORMONAL CONTRACEPTION IN DOGS
26 Surgical Neutering
26.1 OVARIOHYSTERECTOMY OF THE BITCH
26.2 OVARIECTOMY OF THE BITCH
26.3 ORCHIDECTOMY (CASTRATION) OF THE DOG
26.4 VASECTOMY
27 Pharmacological Control of Reproduction
27.1 PROGESTOGENS
27.2 OESTROGENS
27.3 ANDROGENS
27.4 GONADOTROPHINS
27.5 GONADOTROPHIN-RELEASING HORMONE (GnRH) AGONISTS
27.6 GONADOTROPHIN-RELEASING HORMONE (GnRH) ANTAGONISTS
27.7 PROSTAGLANDINS
27.8 PROGESTERONE ANTAGONISTS
27.9 PROLACTIN INHIBITORS
27.10 OXYTOCIN AND ERGOT PREPARATIONS
27.11 SYNTHETIC ANTI-ANDROGENS
28 Reproductive Biotechnology
28.1 ARTIFICIAL INSEMINATION
28.2 ENHANCING FERTILITY IN MALES AND FEMALES WITH REPRODUCTIVE TRACT DISEASE
28.3 HARVESTING GONADAL TISSUE
28.4 CLONING
Index
This edition first published 2013
© 2013 by John Wiley & Sons, Ltd
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Library of Congress Cataloging-in-Publication Data
England, Gary C. W.
Dog breeding, whelping and puppy care / Gary C.W. England.
p. cm.
Includes bibliographical references and index.
ISBN 978-0-470-67313-3 (pbk. : alk. paper) 1. Dogs–Breeding. 2. Dogs–Reproduction. 3. Veterinary obstetrics. I. Title.
SF427.2.E54 2013
636.7'0887–dc23
2012022786
A catalogue record for this book is available from the British Library.
Wiley also publishes its books in a variety of electronic formats. Some content that appears in print may not be available in electronic books.
Cover images: background – iStockphoto/© Trevor Reeves; puppies nursing – iStockphoto/© Nikola Miljkovic; canine sperm – courtesy of Gary England
Cover design by Mark Lee (www.hisandhersdesign.co.uk)
Preface
My endeavour throughout the writing of this book has been to provide a clear and practical guide that is clinically useful for the breeder, veterinary student and veterinary surgeon. I have tried to maintain an easy-to-read style with illustrations that are relevant and explain the subject area and aid understanding of normal and abnormal reproduction as well as common reproductive techniques.
Much of my clinical and research work over the last 25 years has been in the field of canine reproduction and this book represents my view of the subject area. At the suggestion of colleagues the text is written as a monograph and is referenced using my own peer-reviewed publications which demonstrate the impact of my contribution to the field. I hope that my interest and explanations will inspire further study in this exciting discipline.
The work involved in this book is a tribute to the late Dr W.E. Allen, my teacher and friend.
Gary C.W. England
Section 1Breeding, Reproduction and Care of the Normal Animal
1
Care of the Juvenile, Genetics and Planning of Breeding
Puppies are most commonly weaned from their dam at approximately 6 weeks of age and will then be placed into a new home.
the exact time of homing will depend upon the opinion of the breeder and the availability of a new home; early placement at 6 weeks has many advantages relating to behavioural development of the pup, but attention needs to be paid to health care and preventative treatments
whilst pups may arrive in a new home ‘fully weaned’, there are substantial physiological changes in the gastrointestinal tract which are not complete until several months of age; these put the pup at risk of gastrointestinal upset following minor dietary change
there are several important phases of behavioural development, including (1) the neonatal period, (2) the transitional period, and (3) the socialization period (
Figure 1.1
)
on arrival at a new home, pups will have passed through the neonatal and transitional periods (these are discussed later in 15.2) and will be within the socialization period which is probably one of the most important phases of its life
Figure 1.1 Schematic representation of attraction and fear scores for pups demonstrating the ‘critical period’ in relation to the neonatal, transitional and socialization periods.
this period classically starts at 3–4 weeks of age and is complete at approximately 12 weeks
during this time pups should be learning to live as part of the pack (or household)
the period starts with a general acceptance by the pup that anything experienced is not harmful, therefore the pup will not be frightened by exposure to something new; in the wild exposure to new things at an early age would be regulated by the dam and therefore can be considered safe
the period ends with the pup responding to new stimuli with a fear response; in the wild as the pup roams from the nest it is more likely to meet hostile situations and these things should first be treated as dangerous
as the pup moves through the socialization period it starts to learn what is safe, what is dangerous and how to interact with others in the social hierarchy
a ‘critical period’ between 5 and 7 weeks of age has been identified and this is the time when the pup should be exposed to as many stimuli as possible so that the pup will accept these as normal and will not be fearful of them
for appropriate socialization of the pup into the human household it is imperative that the new owner recognizes the importance of the socialization period and provides a rich, stimulating and social environment:
Lesen Sie weiter in der vollständigen Ausgabe!
Lesen Sie weiter in der vollständigen Ausgabe!
Lesen Sie weiter in der vollständigen Ausgabe!
Lesen Sie weiter in der vollständigen Ausgabe!
Lesen Sie weiter in der vollständigen Ausgabe!
Lesen Sie weiter in der vollständigen Ausgabe!
Lesen Sie weiter in der vollständigen Ausgabe!
Lesen Sie weiter in der vollständigen Ausgabe!
Lesen Sie weiter in der vollständigen Ausgabe!
Lesen Sie weiter in der vollständigen Ausgabe!
Lesen Sie weiter in der vollständigen Ausgabe!
Lesen Sie weiter in der vollständigen Ausgabe!
Lesen Sie weiter in der vollständigen Ausgabe!
Lesen Sie weiter in der vollständigen Ausgabe!
Lesen Sie weiter in der vollständigen Ausgabe!
Lesen Sie weiter in der vollständigen Ausgabe!
Lesen Sie weiter in der vollständigen Ausgabe!
Lesen Sie weiter in der vollständigen Ausgabe!
Lesen Sie weiter in der vollständigen Ausgabe!
Lesen Sie weiter in der vollständigen Ausgabe!
Lesen Sie weiter in der vollständigen Ausgabe!
Lesen Sie weiter in der vollständigen Ausgabe!
Lesen Sie weiter in der vollständigen Ausgabe!
Lesen Sie weiter in der vollständigen Ausgabe!
Lesen Sie weiter in der vollständigen Ausgabe!
Lesen Sie weiter in der vollständigen Ausgabe!
Lesen Sie weiter in der vollständigen Ausgabe!
Lesen Sie weiter in der vollständigen Ausgabe!
Lesen Sie weiter in der vollständigen Ausgabe!
Lesen Sie weiter in der vollständigen Ausgabe!
